Turtle Reader

The individuals, and they are the greater number, which inhabit the lower country, can scarcely taste a drop of water throughout the year; but they consume much of the succulent cactus, the branches of which are occasionally broken off by the wind. I several times threw a piece to two or three of them when together; and it was amusing enough to see them trying to seize and carry it away in their mouths, like so many hungry dogs with a bone. They eat very deliberately, but do not chew their food. The little birds are aware how harmless these creatures are: I have seen one of the thick-billed finches picking at one end of a piece of cactus (which is much relished by all the animals of the lower region), whilst a lizard was eating at the other end; and afterwards the little bird with the utmost indifference hopped on the back of the reptile.

I opened the stomachs of several, and found them full of vegetable fibres and leaves of different trees, especially of an acacia. In the upper region they live chiefly on the acid and astringent berries of the guayavita, under which trees I have seen these lizards and the huge tortoises feeding together. To obtain the acacia-leaves they crawl up the low stunted trees; and it is not uncommon to see a pair quietly browsing, whilst seated on a branch several feet above the ground. These lizards, when cooked, yield a white meat, which is liked by those whose stomachs soar above all prejudices. Humboldt has remarked that in intertropical South America all lizards which inhabit dry regions are esteemed delicacies for the table. The inhabitants state that those which inhabit the upper damp parts drink water, but that the others do not, like the tortoises, travel up for it from the lower sterile country. At the time of our visit, the females had within their bodies numerous, large, elongated eggs, which they lay in their burrows: the inhabitants seek them for food.

These two species of Amblyrhynchus agree, as I have already stated, in their general structure, and in many of their habits. Neither have that rapid movement, so characteristic of the genera Lacerta and Iguana. They are both herbivorous, although the kind of vegetation on which they feed is so very different. Mr. Bell has given the name to the genus from the shortness of the snout: indeed, the form of the mouth may almost be compared to that of the tortoise: one is led to suppose that this is an adaptation to their herbivorous appetites. It is very interesting thus to find a well-characterised genus, having its marine and terrestrial species, belonging to so confined a portion of the world. The aquatic species is by far the most remarkable, because it is the only existing lizard which lives on marine vegetable productions. As I at first observed, these islands are not so remarkable for the number of the species of reptiles, as for that of the individuals, when we remember the well-beaten paths made by the thousands of huge tortoises—the many turtles—the great warrens of the terrestrial Amblyrhynchus—and the groups of the marine species basking on the coast-rocks of every island—we must admit that there is no other quarter of the world where this Order replaces the herbivorous mammalia in so extraordinary a manner. The geologist on hearing this will probably refer back in his mind to the Secondary epochs, when lizards, some herbivorous, some carnivorous, and of dimensions comparable only with our existing whales, swarmed on the land and in the sea. It is, therefore, worthy of his observation that this archipelago, instead of possessing a humid climate and rank vegetation, cannot be considered otherwise than extremely arid, and, for an equatorial region, remarkably temperate.

To finish with the zoology: the fifteen kinds of sea-fish which I procured here are all new species; they belong to twelve genera, all widely distributed, with the exception of Prionotus, of which the four previously known species live on the eastern side of America. Of land-shells I collected sixteen kinds (and two marked varieties) of which, with the exception of one Helix found at Tahiti, all are peculiar to this archipelago: a single fresh-water shell (Paludina) is common to Tahiti and Van Diemen’s Land. Mr. Cuming, before our voyage, procured here ninety species of sea-shells, and this does not include several species not yet specifically examined, of Trochus, Turbo, Monodonta, and Nassa. He has been kind enough to give me the following interesting results: of the ninety shells, no less than forty-seven are unknown elsewhere—a wonderful fact, considering how widely distributed sea-shells generally are. Of the forty-three shells found in other parts of the world, twenty-five inhabit the western coast of America, and of these eight are distinguishable as varieties; the remaining eighteen (including one variety) were found by Mr. Cuming in the Low Archipelago, and some of them also at the Philippines. This fact of shells from islands in the central parts of the Pacific occurring here, deserves notice, for not one single sea-shell is known to be common to the islands of that ocean and to the west coast of America. The space of open sea running north and south off the west coast separates two quite distinct conchological provinces; but at the Galapagos Archipelago we have a halting-place, where many new forms have been created, and whither these two great conchological provinces have each sent several colonists. The American province has also sent here representative species; for there is a Galapageian species of Monoceros, a genus only found on the west coast of America; and there are Galapageian species of Fissurella and Cancellaria, genera common on the west coast, but not found (as I am informed by Mr. Cuming) in the central islands of the Pacific. On the other hand, there are Galapageian species of Oniscia and Stylifer, genera common to the West Indies and to the Chinese and Indian seas, but not found either on the west coast of America or in the central Pacific. I may here add, that after the comparison by Messrs. Cuming and Hinds of about 2000 shells from the eastern and western coasts of America, only one single shell was found in common, namely, the Purpura patula, which inhabits the West Indies, the coast of Panama, and the Galapagos. We have, therefore, in this quarter of the world, three great conchological sea-provinces, quite distinct, though surprisingly near each other, being separated by long north and south spaces either of land or of open sea.

I took great pains in collecting the insects, but excepting Tierra del Fuego, I never saw in this respect so poor a country. Even in the upper and damp region I procured very few, excepting some minute Diptera and Hymenoptera, mostly of common mundane forms. As before remarked, the insects, for a tropical region, are of very small size and dull colours. Of beetles I collected twenty-five species (excluding a Dermestes and Corynetes imported wherever a ship touches); of these, two belong to the Harpalidæ, two to the Hydrophilidæ, nine to three families of the Heteromera, and the remaining twelve to as many different families. This circumstance of insects (and I may add plants), where few in number, belonging to many different families, is, I believe, very general. Mr. Waterhouse, who has published153 an account of the insects of this archipelago, and to whom I am indebted for the above details, informs me that there are several new genera; and that of the genera not new, one or two are American, and the rest of mundane distribution. With the exception of a wood-feeding Apate, and of one or probably two water-beetles from the American continent, all the species appear to be new.

The botany of this group is fully as interesting as the zoology. Dr. J. Hooker will soon publish in the “Linnean Transactions” a full account of the Flora, and I am much indebted to him for the following details. Of flowering plants there are, as far as at present is known, 185 species, and 40 cryptogamic species, making together 225; of this number I was fortunate enough to bring home 193. Of the flowering plants, 100 are new species, and are probably confined to this archipelago. Dr. Hooker conceives that, of the plants not so confined, at least 10 species found near the cultivated ground at Charles Island have been imported. It is, I think, surprising that more American species have not been introduced naturally, considering that the distance is only between 500 and 600 miles from the continent, and that (according to Collnet, page 58) drift-wood, bamboos, canes, and the nuts of a palm, are often washed on the south-eastern shores. The proportion of 100 flowering plants out of 185 (or 175 excluding the imported weeds) being new, is sufficient, I conceive, to make the Galapagos Archipelago a distinct botanical province; but this Flora is not nearly so peculiar as that of St. Helena, nor, as I am informed by Dr. Hooker, of Juan Fernandez. The peculiarity of the Galapageian Flora is best shown in certain families;—thus there are 21 species of Compositæ, of which 20 are peculiar to this archipelago; these belong to twelve genera, and of these genera no less than ten are confined to the archipelago! Dr. Hooker informs me that the Flora has an undoubted Western American character; nor can he detect in it any affinity with that of the Pacific. If, therefore, we except the eighteen marine, the one fresh-water, and one land-shell, which have apparently come here as colonists from the central islands of the Pacific, and likewise the one distinct Pacific species of the Galapageian group of finches, we see that this archipelago, though standing in the Pacific Ocean, is zoologically part of America.